M. Sadoud1*, J.F. Hocquette2 1Hassiba Benbouali Chlef University, Faculty of Science, Nature and Life, Department of Biology, (02000), Chlef, Algeria 2INRAE, Clermont Auvergne University, Vetagro Sup, UMR1213, Herbivore Research Unit, Theix 63122 Saint-Genès-Champanelle, France (Manus